![]() These measurements have revealed that the pH mito is in dynamic equilibrium with the cytosolic pH and that spontaneous pH mito elevations coinciding with ΔΨ m drops occur in single mitochondria. In this Perspective, we discuss the putative roles of the pH mito and review the different techniques used to measure pH mito and ΔpH m in isolated mitochondria and in intact cells, focusing on our recent results obtained with genetically encoded pH-sensitive indicators. ΔpH m, however, drives the fluxes of metabolic substrates required for mitochondrial respiration and the activity of electroneutral ion exchangers that maintain mitochondria osmolarity and volume, and recent studies indicate that the mitochondrial pH (pH mito) plays an important and underappreciated role in physiological and pathological situations such as apoptosis, neurotransmission, and insulin secretion. Because ΔΨ m contributes most of the Δp and can be easily measured in intact cells with fluorescent dyes, most studies ignore the contribution of ΔpH m and only record changes in ΔΨ m to track the metabolic state of mitochondria. Studies in isolated mitochondria have established that the chemical gradient for protons (ΔpH m) and the mitochondrial membrane potential (ΔΨ m) contribute independently to the proton-motive force (Δp) that drives the synthesis of ATP. The generation of a proton gradient across the inner mitochondrial membrane (IMM) is an essential energy conservation event that couples the oxidation of carbohydrates and fat to the synthesis of ATP. ![]()
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![]() ![]() We became a society that was not only modern and wealthy, but dignified and humane.” “We didn’t just change the infrastructure. With reports spanning from the ghostly Great Barrier Reef, to the annual smoke-choked skies of the Pacific Northwest, to post-hurricane Puerto Rico, to a Vatican attempting an unprecedented “ecological conversion,” Klein makes the case that we will rise to the existential challenge of climate change only if we are willing to transform the systems that produced this crisis.Īn expansive, far-ranging exploration that sees the battle for a greener world as indistinguishable from the fight for our lives, On Fire captures the burning urgency of the climate crisis, as well as the fiery energy of a rising political movement demanding a catalytic Green New Deal. Delving into topics ranging from the clash between ecological time and our culture of “perpetual now,” to the soaring history of humans changing and evolving rapidly in the face of grave threats, to rising white supremacy and fortressed borders as a form of “climate barbarism,” this is a rousing call to action for a planet on the brink. These long-form essays show Klein at her most prophetic and philosophical, investigating the climate crisis not only as a profound political challenge but as a spiritual and imaginative one, as well. On Fire: The (Burning) Case for a Green New Deal gathers for the first time more than a decade of her impassioned writing, and pairs it with new material on the staggeringly high stakes of our immediate political and economic choices. In lucid, elegant dispatches from the frontlines of contemporary natural disaster, she pens surging, indispensable essays for a wide public: prescient advisories and dire warnings of what future awaits us if we refuse to act, as well as hopeful glimpses of a far better future.
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